They are the exact same bike mechanically with these changes to create Z400:
- no fully-coverage bodywork, a naked sport bike
- no windshield
- slightly softer suspension
- taller handlebars
- mirrors mounted on handlebars
Basically a more "standard" bike like old UJM versus a "sport" bike. More comfortable due to more upright seating position. Although some people complain about seeing a lot of their elbows in mirrors. A lot of naked bike users end up putting windshields on later to reduce wind noise and buffeting at at highway speeds.
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